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It is plausible to postulate that the first and second GSH molecule should interact during the reductive half-reaction of the dithiol mechanism with the 1st and 2nd glutathione-interaction site, respectively

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Kubera M, Obuchowicz E, Goehler L et al (2011) In animal models, psychosocial stress-induced (neuro)inflammation, apoptosis and reduced neurogenesis are associated to the onset of depression

This is why MTHFR mutations are often seen as the most problematic of the methylation mutations - because the entire system hinges on having enough MTHFR

It has biological relevance to tissue remodeling